Latest publications:

Martín-Fernández C, Solti Á, Czech V, Kovács K, Fodor F, Gárate A, Hernández-Apaolaza L, Lucena J J (2017) Response of soybean plants to the application of synthetic and biodegradable Fe chelates and Fe complexes. Plant Physiology and Biochemistry 118: 579-588.

Rév A, Tóth B, Solti Á, Sipos Gy, Fodor F (2017) Responses of Szarvasi-1 energy grass to sewage sludge treatments in hydroponics. Plant Physiology and Biochemistry 118: 627-633.


Fodor F. (ed.) 15th International Symposium on Iron Nutrition and Interactions in Plants, Programme and Book of Abstracts (2010)

Guest editor of the PPB Special issue: 15th International Symposium on Iron Nutrition and Interactions in Plants (Budapest, Hungary, 2010)
Shortcut to the issue: Volume 49, Issue 5, Pages 461-564 (May 2011)

Fodor F. (ed.) A növényi anyagcsere élettana (Physiology of Plant Metabolism) (2013) e-book, ELTE  pp.367

Fodor F. (ed.) Paál Árpádtól a Molekuláris Növénybiológiáig. Tudományos ülés Paál Árpád születésének 125. és intézetigazgatói kinevezésének 85. évfordulója alkalmából. (From Árpád Paál to the Molecular Plant Biology. Scientific session on the 125th anniversary of Árpád Paál’s birth and 85th anniversary of his nomination as director.)(2014) conference proceedings, Növényélettani és Molekuláris Növénybiológiai Tanszék, TTK, ELTE pp. 70


  1. Böddi B., Cseh E., Fodor F., Király I., Nyitrai P., Sárvári É., Szabó E., Szigeti Z., Tamás L. (1994) Növényélettani gyakorlatok (Plant Physiology Practices) (Ed. Láng F.) p. 236., ELTE Kiadó, Budapest.

  2. Böddi B., Cseh E., Fodor F., Király I., Láng F., Lásztity D., Nyitrai P., Sárvári É., Szabó E., Szigeti Z. (1999) Növényélettan: A növényi anyagcsere. (Plant Physiology: The Plant Metabolism) (Ed. Láng F.) pp. 998, ELTE Kiadó, Budapest.

  3. Bratek Z., Fodor F., Király I., Nyitrai P., Parádi I., Sárvári É., Solti Á., Szabó E., Szigeti Z., Tamás L. (2013) A növényi anyagcsere élettana. (Physiology of Plant Metabolism), e-book (Ed. Fodor F.) pp.367

  4. Fodor F., Nyitrai P., Oszvald M., Rudnóy Sz., Sárvári É., Solti Á., Szigeti Z., Tamás L., Tóth G. (2013) Növényélettani vizsgáló módszerek. (Research methods in Plant Physiology) e-book (Ed. Tamás L.) pp.367

  5. Bratek Z., Fodor F., Rudnóy Sz., Solti Á., Tamás L. (2016) Research Methods in Plant Biology. angol nyelvű elektronikus jegyzet (Szerk. Tamás L.) pp.367

Book chapters:

  1. Fodor F. (2002) Physiological responses of vascular plants to heavy metals. in eds. M.N.V. Prasad and K. Strzalka, Physiology and Biochemistry of Metal Toxicity and Tolerance in Plants, 149-177. Kluwer Academic Publishers, Dordrecht.

  2. Fodor F. (2006) Heavy metals competing with iron under conditions involving phytoremediation. In eds. L.L.Barton and J.Abadía, Iron Nutrition in Plants and Rhizospheric Microorganisms, 129-151., Springer.

  3. Tóth B., Fodor F. (2015) Alternative ways of utilization of wastes. in eds. R. Efe, C. Bizzarri, I. Cürebal and G. N. Nyusupova, Environment and Ecology at the Beginning of the 21th Century, 11-27. St. Kliment Ohridski University Press, Sofia

Full papers:

  1. Fodor F., Cseh E. (1993) Effect of different nitrogen-forms and iron-chelates on the development of stinging nettle. J. Plant Nutr. 16: 2239-2253.

  2. Fodor F, Böddi B., Sárvári É., Záray Gy., Cseh E., Láng F. (1995) Correlation of iron content, spectral forms of chlorophyll and chlorophyll-proteins in iron deficient cucumber (Cucumis sativus). Phys. Plant. 93: 750-756.

  3. Dao Thi Phuong D., Tatár E., Varga I., Záray Gy., Cseh E., Fodor F. (1995) Accumulation and translocation of lead in cucumber plants monitored by Graphite Furnace-Atomic Absorption Spectometry. Microchem. J. 51: 145-150.

  4. Záray Gy., Dao Thi Phuong D., Varga I., Varga A., Kántor T., Cseh E., Fodor F. (1995) Influences of lead contamination and complexing agents on the metal uptake of cucumber. Microchem. J. 51: 207-213.

  5. Babos K., Pócs T., Fodor F. (1995) Xylotomy of two endemic senecio species from Kenya. Acta Botanica Hungarica 39: 235-242.

  6. Fodor F., Cseh E., Dao Thi Phuong D., Záray Gy. (1996) Uptake and mobility of lead in Urtica dioica L. Acta Phytogeographica Suecica 81: 106-108.

  7. Fodor F., Sárvári É., Láng F., Szigeti Z., Cseh E. (1996) Effects of Pb and Cd on cucumber depending on the Fe-complex in the culture solution. J. Plant Physiol. 148: 434-439.

  8. Záray Gy., Varga A., Fodor F., Cseh E. (1997) Microanalytical investigation of xylem sap of cucumber by Total Reflection X-Ray Fluorescence Spectroscopy. Microchem. J. 55: 64-71.

  9. Varga A., Záray Gy., Fodor F., Cseh E. (1997) Study of interaction of iron and lead during their uptake process in wheat roots by Total Reflection X-Ray Fluorescence Spectrometry. Spectrochimica Acta Part B 52, 1027-1032.

  10. Fodor F., Cseh E., Varga A., ZarayGy. (1998) Lead uptake, distribution and remobilization in cucumber. J. Plant Nutr. 21: 1363-1373.

  11. Tatár E., Mihucz V.G., Varga A., Záray Gy., Fodor F. (1998) Determination of Organic Acids in Xylem Sap of Cucumber: Effect of Lead Contamination. Microchem. J. 58: 306-314.

  12. Sárvári É., Fodor F., Cseh E., Varga A., Záray Gy., Zolla L. (1999) Relationship between changes in ion content of leaves and thylakoid composition in cucumber under Cd and Pb stress. Z. Naturforsch. Part C 54, 746-753.

  13. Varga A., Martínez R.M.G., Záray Gy., Fodor F. (1999) Investigation of effects of cadmium, lead, nickel and vanadium contamination on the uptake and transport processes in cucumber plants by TXRF-spectrometry, Spectrochim. Acta B 54, 1455-1462.

  14. Cseh E., Fodor F., Varga A., Záray Gy. (2000) Effect of lead treatment on the distribution of essential elements in cucumber. J. Plant Nutr. 23: 1095-1105.

  15. Varga A., Záray Gy., Fodor F. (2000) Investigation of element distributions between the symplasm and apoplasm of cucumber plants by TXRF spectrometry. Microchem. J. 67: 257-264.

  16. Tatar E., Mihucz V.G., Kmethy B., Zaray G., Fodor F. (2000) Determination of organic acids and their role in nickel transport within cucumber plants. Microchem J. 67: 73-81.

  17. Sárvári É., Gáspár L., Fodor F., Cseh E., Kröpfl K., Varga A., Baron M. (2002) Comparison of the effects of Pb treatment on thylakoid development in poplar and cucumber plants. Proceedings of the 7th Hungarian Congress on Plant Physiology, Acta Biologica Szegediensis 46: 163-165.

  18. Lenti K., Fodor F., Böddi B. (2002) Mercury inhibits the activity of the NADPH:protochlorophyllide oxidoreductase (POR). Photosynthetica 40: 145-151.

  19. Varga A., Záray Gy., Fodor F. (2002) Determination of element distribution between  apoplasm and symplasm of cucumber plant parts by total reflection X-ray fluorescence spectrometry. J. Inorganic Biochemistry 1989: 149-154.

  20. Fodor F., Gáspár L., Morales F., Gogorcena Y., Lucena J.J., Cseh E., Kröpfl K., Abadía J., Sárvári É. (2005) Effect of two iron sources on iron and cadmium allocation in poplar (Populus alba) plants exposed to cadmium. Tree Phys. 25: 1173-1180.

  21. Gáspár L., Czövek P., Fodor F., Hoffmann B., Nyitrai P., Király I., Sárvári É. (2005) Greenhouse testing of new wheat cultivars compared to those with known drought tolerance. Proceedings of the 8th Hungarian Congress of Plant Physiology, Acta Biologica Szegediensis 49: 97-98.

  22. Kovács K., Kuzmann E., Fodor F., Vértes A., Kamnev A.A. (2005) Mössbauer study of iron uptake in cucumber root. Hyperfine Interact. 165: 289-294.

  23. Mihucz VG, Tatar E, Virag I, Cseh E, Fodor F, Zaray G (2005) Arsenic speciation in xylem sap of cucumber (Cucumis sativus L.) Anal. Bioanal. Chem. 383: 461-466.

  24. Czech V, Czövek P, Fodor J, Bóka K, Fodor F, Cseh E (2008) Investigation of arsenate phytotoxicity in cucumber plants. Acta Biologica Szegediensis 52: 79-80.

  25. Kovács K, Kuzmann E, Fodor F, Cseh E, Homonnay Z, Vértes A (2008) Mössbauer investigation of iron uptake in wheat. Hyperfine Interact. 185: 63-67.

  26. Kovács K, Kuzmann E, Tatár E, Vértes A, Fodor F (2009) Investigation of iron pools in cucumber roots by Mössbauer spectroscopy: direct evidence for the Strategy I iron uptake mechanism. Planta 229: 271-278.

  27. Lévai L, Fodor F, Marozsán M (2009) Effects of biotic stress on the uptake and distribution of some elements. Cereal Research Communications DOI:10.1556/CRC.37.2009.Suppl.2

  28. Kovács K, Kuzmann E, Vértes A, Lévai L, Cseh E, Fodor F (2010) Effect of cadmium on iron uptake in cucumber roots: A Mössbauer-spectroscopic study. Plant and Soil 327:49-56.

  29. Gyuricza V, Fodor F, Szigeti Sz (2009) Phytotoxic effects of heavy metal contaminated soil reveal limitations of extract-based ecotoxicological tests Water, Air and Soil Pollution 210:113-122.

  30. Rabnecz Gy, Záray Gy, Lévai L, Fodor F (2009) Effect of heavy metals on the leaf disc ferricyanide reduction in cucumber. Acta Agronomica Hungarica, 57: 307-320.

  31. Vashegyi I, Cseh E, Lévai L, Fodor F (2011) Chelator-enhanced lead uptake, accumulation in energy grass (Agropyron elongatum cv. "Szarvasi-1"). International Journal of Phytoremediation, 13:302-315.

  32. Sárvári É, Gáspár L, Solti Á, Mészáros I, Záray Gy, Fodor F (2010) Cd-Fe Interactions: comparison of the effects of iron deficiency and cadmium on growth and phtotosynthetic performance in poplar. Acta Biologica Hungarica 61 (suppl.): 136-148.

  33. Czech V, Cseh E, Fodor F (2011) Arsenate induces water stress. Journal of Plant Nutrition, 34: 1–11.

  34. Solti Á., Sárvári É., Tóth B., Basa B., Lévai L., Fodor F. (2011) Cd affects the translocation of some metals either Fe-like or Ca-like way in Poplar. Plant Physiology and Biochemistry 49: 494-498.

  35. Sárvári É., Solti Á., Basa B., Mészáros I., Lévai L., Fodor F. (2011) Impact of moderate Fe excess under Cd stress on the photosynthetic performance of poplar (Populus jacquemontiana var. glauca cv. Kopeczkii) Plant Physiology and Biochemistry 49: 499-505.

  36. Csog Á., Mihucz V.G., Tatár E., Fodor F., Virág I., Majdik C., Záray Gy. (2011) Accumulation and distribution of iron, cadmium, lead and nickel in cucumber plants grown in hydroponics containing two different chelated iron supplies. Journal of Plant Physiology 168:1038-1044.

  37. Solti Á, Szöllősi E, Gémes Juhász A, Mészáros I, Fodor F, Sárvári É (2011) Significance of antioxidative defence under long-term Cd stress. Acta Biologica Szegediensis 55:151-153.

  38. Solti Á, Gáspár L, Vági P, Záray Gy, Fodor F, Sárvári É (2011) Cd, Fe and light sensitivity: interrelationships in Cd treated Populus. OMICS - A Journal of Integrative Biology, 15: 811-818.

  39. Solti Á, Kovács K, Basa B, Vértes A, Sárvári É, Fodor F (2012) Uptake and incorporation of iron in sugar beet chloroplasts. Plant Physiol. Biochem. 52: 91-97.

  40. Mihucz V G, Csog Á, Fodor F, Tatár E, Szoboszlai N¸ Silaghi-Dumitrescu L, Záray G. (2012) Impact of two iron(III) chelators on the iron, cadmium, lead and nickel accumulation in poplar grown under heavy metal stress in hydroponics. J. Plant Physiology 169: 561-566.

  41. Tóth B, Solti Á, Fodor F, Bóka K, Veres Sz (2012): Green development: waste water or green water? Növénytermelés/Crop production 61. Suppl.: 467-470. (ISSN 0546-8191)

  42. Carrasco J; Kovács K; Czech V; Fodor F; Lucena JJ; Vértes A; Hernández-Apaolaza L (2012): Influence of pH, iron source and Fe/ligand ratio on the iron speciation in lignosulfonate complexes by using Mössbauer spectroscopy. Implications on their fertilizer properties. Journal of Agricultural and Food Chemistry, 60: 3331-3340.

  43. Fodor F; Kovács K; Czech V; Solti Á; Tóth B; Lévai L; Bóka K; Vértes A (2012): Effects of short term iron citrate treatments at different pH values on roots of iron deficient cucumber: a Mössbauer analysis. Journal of Plant Physiology 169: 1615-1622.

  44. Yunta F, Di Foggia M, Bellido V, Morales M, Tessarin P, López-Rayo S, Tinti A, Kovács K, Klencsár Z, Fodor F, Rombola A D (2013) Blood meal-based compound. Good choice as iron fertilizer for organic farming. Journal of Agricultural and Food Chemistry 61: 3995-4003.

  45. Sipos Gy, Solti Á, Czech V, Vashegyi I, Tóth B, Cseh E, Fodor F (2013) Heavy metal accumulation and tolerance of energy grass (Elymus elongatus subsp. ponticus cv. Szarvasi-1) grown in hydroponic culture. Plant Physiology and Biochemistry 68: 96-103.

  46. Tóth B, Nagy GL, Nagy L, Lévai L, Fodor F, Solti Á, Veres Sz (2013) Possible recycling of industrial wastes and by-products in agriculture. Procedia Environmental Sciences 18: 737-741.

  47. Kovács K, Czech V, Fodor F, Solti Á, Lucena JJ, Santos-Rosell S, Hernández-Apaolaza L. (2013) Characterization of Fe-leonardite complexes, as novel natural iron fertilizers. Journal of Agricultural and Food Chemistry 61: 12200-12210

  48. Basa B, Lattanzio G, Solti Á, Tóth B, Abadía J, Fodor F, Sárvári É. (2014) Changes induced by cadmium stress and iron deficiency in the composition and organization of thylakoid complexes in sugar beet (Beta vulgaris L.). Environmental and Experimental Botany 101:1-11.

  49. Solti Á., Müller B., Czech V., Sárvári É., Fodor F. (2014) Functional characterization of the chloroplast ferric chelate oxidoreductase enzyme. New Phytologist 202:920-928.

  50. Kovács K, Pechoušek J, Machala L, Zbořil R, Klencsár Z, Solti Á, Tóth B, Müller B, Pham H D., Kristóf Z, Fodor F. (2016) Revisiting the iron pools in cucumber roots: identification and localization. Planta 244: 167-179.

  51. Solti Á., Sárvári É, Tóth B, Mészárosc I, Fodor F (2016) Incorporation of iron into chloroplasts triggers the restoration of cadmium induced inhibition of photosynthesis. Journal of Plant Physiology. 202:97-106.

  52. Homonnay Z, Tolnai Gy, Fodor F, Solti Á, Kovács K, Kuzmann E, Ábrahám A, Szabó E Gy, Németh P, Szabó L, Klencsár Z.(2016) Iron oxide nanoparticles for plant nutrition? A preliminary Mössbauer study. Hyperfine Interactions 237:127.

  53. Solti Á, Sárvári É, Szöllősi E, Tóth B, Mészáros I, Fodor F, Szigeti Z. (2016) Stress hardening under long-term cadmium treatment depends on the activation of antioxidative defence and the iron acquisition of chloroplasts in Populus. Zeitshrift für Naturforschung C-A Journal of Biosciences 71: 323-334.

  54. Solti Á, Kovács K, Müller B, Vázquez S, Hamar É, Pham H D, Tóth B, Abadía J, Fodor F. (2016) Does a voltage-sensitive outer envelope transport mechanismcontributes to the chloroplast iron uptake? Planta 244: 1303–1313.

  55. Gerényi A, Czech V, Fodor F, Vincze L, Szalóki I. (2016) In-vivo XANES measuring techniques for studying the arsenic uptake in cucumber plant. X-Ray Spectrometry 46: 143-150.

Full papers in Hungarian:

  1. Cseh E., Fodor F. (1993) A vasellátotság és a N-formák együttes hatása az uborka növekedésére és klorofill-szintézisére. Bot. Közl. 80: 19-29.

  2. Cseh E., Fodor F. (1994) Megszüntethető-e az élettani vashiány? A N-formák és a pH hatása a vas mobilitására. Bot. Közl. 81: 21-27.

  3. Cseh E., Váradi Gy., Fodor F. (1994) Fe-komplexek és N-formák hatása az uborka Fe-abszorpciójára, -felvételére és -transzlokációjára. Bot. Közl. 81: 47-55.

  4. Cseh E., Váradi Gy., Fodor F.(1994) A N-formák hatása az uborka Fe-felvételére és transzlokációjára HCO3- jelenlétében. Bot. Közl. 81: 195-199.

  5. Fodor F. (2003) Ólom- és kadmiumstressz növényekben. Bot Közl. 90: 107-120.

Fullpapers in conference proceedings books:

  1. Láng F., Sárvári É., Szigeti Z., Fodor F., Cseh E. (1995) Effects of heavy metals on the photosynthetic apparatus in cucumber. (In the proceedings of the Xth International Photosynthesis Congress, Montpellier, France.) P. Mathis (ed.), Photosynthesis: from Light to Biosphere, Vol. IV, pp. 533-536, Kluwer Academic Publishers.

  2. Cseh E., Fodor F. (1997) Role of boron in the plasmalemma turbo reductase activity. In the proceedings of the International Symposium on Boron in Soils and Plants, Chiang Mai, Thailand. R.W. Bell and B. Rerkasem (eds.) Boron in Soils and Plants. 175-177, Kluwer Academic Publishers.

  3. Fodor F., Cseh E., Kovács-Láng E. (1998) Potassium uptake and allocation of Bothriochloa ischaemum. In the proceedings of the International Society of Root Research 5th Symposium, Clemson, South Carolina, USA. pp. 533-540, Kluwer Academic Publishers.

  4. Láng F., Szigeti Z., Fodor F., Cseh E., Zolla L., Sárvári É. (1998) Influence of Cd and Pb on the ion content, growth and photosynthesis in cucumber. In the proceedings of the XIth International Congress on Photosynthesis, Budapest. pp 2693-2697.

  5. Láng F., Sárvári É., Gáspár L., Fodor F., Cseh E. (2001) Influence of light intensity on thylakoid development under Cd stress in poplar. In the Proceedings of the 12th International Congress on Photosynthesis, Brisbane, S3-023.

  6. Sárvári É., Szigeti Z., Fodor F., Cseh E., Tussor K., Záray Gy., Veres Sz., Mészáros I. (2001) Relationship of iron deficiency and the altered thylakoid develeopment in Cd treated poplar plants. In the Proceedings of the 12th International Congress on Photosynthesis, Brisbane, S3-025.

  7. Záray Gy., Kröpfl K., Mihucz V.G., Yao J., Cseh E., Fodor F. (2004) Uptake and translocation of arsenic in cucumber plants grown in nutrient solutions contaminated with As(V) or As(III). in Metal Ions in Biology and Medicine, Vol. 8. Eds: MA Cser, I Sziklai László, JC Etienne, Y Maymard, JA Centeno, L Khassanova, Ph Collery, Budapest. p. 186-189.

  8. Mihucz V.G., Kröpfl K., Tatár E., Yao J., Fodor F., Cseh E., Záray Gy. (2004) As(III) és As(V) felvételének vizsgálata uborkanövényekben totál reflexiós röntgenfluoreszcens spektrometriai módszerrel. 47. Magyar Spektrokémiai Vándorgyulés, Szerk: Adányiné Kisbocskói Nóra, Mihucz Viktor Gábor, Balatonföldvár, 202-205.

  9. Tóth B., Veres Sz., Fodor F., Solti Á., Lévai L. (2011): Waste management – Reuse in agriculture? Progress in Environmental Science and Technology, Vol III. (eds. Li S., Wang W., Niu P., An Y.) ISBN978-7-03-030935-8, Proceedings of the 2011 International Sympsium on Environmental Science and Technology, Dongguan, Guangdong Province, China, June 1-4, pp. 945-949.

  10. Fodor F., Cseh E., Solti Á., Czech V., Kovács K. Vaskelátok és vaskomplexek a vashiány orvoslásában: régi probléma, új ismeretek. (Fe-chelates and Fe-complexes in correcting Fe deficiency: old problem, new information.) in Hat évtized az agrár felsőoktatásért. Eds. Jávor A., Fürjné Rádi K., Veres Sz., Debreceni Egyetem Agrár- és Gazdálkodástudományok Centruma, ISBN 978-615-5183-97-3, pp49-55.

  11. Tóth B., Fodor F., Lévai L., Kovács B. (2015): Role of bacteria in Al-toxicity. In: Proceedings of the 9th International Symposium on Plant-Soil Interactions at Low pH (eds: Z. Loncaric, L. Kochian), pp. 14-15. ISBN 978-953-7871-40-6

Abstracts in scientific journals:

  1. Fodor F., Cseh E., Dao Thi Phuong D., Záray Gy. (1994) Effect of chelators on lead, iron and potassium uptake and interactions. Biologia Plantarium 36 suppl. S189.

  2. Láng F., Sárvári É., Szigeti Z., Fodor F., Cseh E. (1996) Changes of the photosynthetic properties of excised cucumber (Cucumis sativus L.) leaves during formation of new roots. Plant Physiology and Biochemistry Spec. Issue, S09-14, p109.

  3. Nyitrai P., Sárvári É., Keresztes Á., Fodor F., Cseh E., Láng F. (2000) Development of thylakoids under the effect of Cd and Pb in cucumber. Plant Physiology and Biochemistry Spec. Issue, S11-12, p104.

  4. Sárvári É., Fodor F., Cseh E., Szigeti Z., Láng F. (2000) Comparison of the effects of Cd stress and Fe-deficiency on the thylakoid development in poplar offsprings. Plant Physiology and Biochemistry Spec. Issue, S19-25, p180.

  5. Fodor F., Cseh E. (2000) Effect of heavy metals on the ferricyanide reductase activity of leaf tissues. Plant Physiology and Biochemistry Spec. Issue, S19-39, p185.

  6. Sárvári É., Gáspár L., Kröpfl K., Fodor F., Cseh E. (2004) Comparison of the effects of Cd toxicity with iron and manganase deficiency on thylakoid development in poplar. Acta Physiologiae Plantarum Spec. Issue, 26: Suppl. 3., AS-063, p210.

  7. Czech V, Czoevek P, Fodor J, Boka K, Fodor F, Cseh E et al. (2008) Investigation of arsenate phytotoxicity in cucumber plants. Acta Biologica Szegediensis Spec. Issue 52: 79-80.

  8. Müller B, Kovács K, Halász K, Kavak Y, Gyuris B, Fodor F, Sárvári É, Solti Á (2017) The chemical forms of Fe used as in vivo substrate in uptake process of chloroplasts. J. Plant Physiology & Pathology 5: Paper 10.4172/2329-955X-C1-012. (3rd Global Summit on Plant Science. Róma, Olaszország: 2017.08.07 -2017.08.09.)

Abstracts in scientific journals in Hungarian:

  1. Cseh E., Fodor F. (1991) A N-formák hatása a vas felvételére és transzlokációjára. Bot. Közl. 78: 192.

  2. Fodor F., Cseh E. (1991) A nagy csalán (Urtica dioica) vashasznosításának vizsgálata. Bot. Közl. 78: 194.


A nitrogén-formák és a komplexképzők hatása a vas és az ólom felvételére és transzlokációjára. (1996) Egyetemi doktori/PhD értekezés, ELTE, Budapest.
(The effect of nitrogen forms and chelating agents on the uptake and translocation of iron and lead, Doctoral/PhD thesis, Eötvös University, Budapest, Hungary)